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Investigations in the amplitude of sounded piano tones.

The relationship between final hammer velocity and maximum amplitude of radiated piano sound was investigated. Piano tones with varying hammer velocities were produced by a computer-monitored acoustic piano containing optical sensors and solenoids, and the sounded tones were recorded and digitized for analysis. Maximum amplitudes over the duration of the sounded tones were linearly proportional to piano hammer velocities for a range of frequencies and hammer velocities. Changes in room acoustics did not alter the linear relationship. Measurements of maximum amplitudes of individual tones and combined tones (dyads) also indicated a linear relationship between the sum of the maximum amplitudes of the individual tones and the maximum amplitude of the dyads. These findings indicate that the principle of superposition holds for peak amplitudes of sounded piano tones. Findings are discussed with regard to production and perception of musical dynamics.

Computer Graphics↗

The dominant role of low-frequency interaural time differences in sound localization.

Two experiments are described in which listeners judge the apparent directions of virtual sound sources-headphone-presented sounds that are processed in order to simulate free-field sounds. Previous results suggest that when the cues to sound direction are preserved by the simulation, the apparent directions of virtual sources are nearly the same as the apparent directions of real free-field sources. In the experiments reported here, the interaural phase relations in the processing algorithms are manipulated in order to produce stimuli in which the interaural time difference cues signal one direction and interaural intensity and pinna cues signal another direction. The apparent directions of these conflicting cue stimuli almost always follow the interaural time cue, as long as the wideband stimuli include low frequencies. With low frequencies removed from the stimuli, the dominance of interaural time difference disappears, and apparent direction is determined primarily by interaural intensity difference and pinna cues.

Adult↗

Neural derivation of sound source location: resolution of spatial ambiguities in binaural cues.

Cues for sound localization are inherently spatially ambiguous. Nevertheless, most neurons in the barn owl's optic tectum (superior colliculus) have receptive fields for broadband noise stimuli that are restricted to a single region of space. This study characterizes the spatial ambiguities associated with two important sets of localization cues, interaural level differences (ILDs) and interaural phase differences (IPDs), and describes how information is integrated within and across frequencies to resolve these ambiguities. The auditory receptive fields of neurons in the optic tectum were measured with free-field sounds presented from a movable loudspeaker. In contrast to the single regions typical for broadband receptive fields, receptive fields for tonal stimuli usually included additional discrete regions of space (accessory fields). Based on acoustic measurements of ILD and IPD cues made in the external ear canals, it was shown that accessory fields corresponded to locations from which sound sources produced ILD and IPD values that were approximately the same as those arising from the broadband receptive field. In addition, accessory fields had inhibitory surrounds, corresponding to locations from which sound sources produced substantially different combinations of ILD and IPD values. Where an accessory field for one frequency overlapped with the inhibitory surround of a second frequency, an excitatory response to the first frequency could be reduced or eliminated by addition of the second frequency. Because tonal receptive fields for different frequencies always overlapped in the region of the broadband receptive field but tended not to overlap elsewhere, this integration of excitation and inhibition can account for the restriction of broadband receptive fields to a single region of space.

Animals↗

Analysis of acoustic elements and syntax in communication sounds emitted by mustached bats.

Mustached bats, Pteronotus parnellii parnellii spend most of their lives in the dark and use their auditory system for acoustic communication as well as echolocation. The sound spectrograms of their communication sounds or "calls" revealed that this species produces a rich variety of calls. These calls consist of one or more of the 33 different types of discrete sounds or "syllables" that are emitted singly and/or in combination. These syllables can be further classified as 19 simple syllables, 14 composites, and three subsyllables. Simple syllables consist of characteristic geometric patterns of CF (constant frequency), FM (frequency modulation), and NB (noise burst) sounds that are defined quantitatively using statistical criteria. Composites consist of simple syllables or subsyllables conjoined without any silent interval. Most syllable types exhibit a large intrinsic variation in their physical structure compared to the stereotypic echolocation pulses. Syllable domains are defined on the basis of multiple parameters, although these can be collapsed onto three dimensions that capture 99% of the measured variation among different types of syllables. Temporal analysis of multisyllabic constructs reveals several syntactical rules for syllable transitions.

Acoustics↗

Spectral cues for sound localization in cats: effects of frequency domain on minimum audible angles in the median and horizontal planes.

Rice et al. [Hear. Res. 58, 132-152 (1992)] classified directional properties of the cat's head-related transfer function (HRTF) into three frequency domains. Low frequencies (< 5 kHz) display a broad azimuth-sensitive spectral peak that establishes interaural level differences, mid frequencies (5-18 kHz) are marked with a single deep spectral notch that changes in frequency as a function of both azimuth and elevation, and high frequencies (18-50 kHz) exhibit a complex pattern of peaks and notches that shows extensive but less systematic changes with sound location. Spectral cues conveyed by the mid frequencies of broadband sounds are important in tasks that require cats to identify the actual location of acoustic stimuli [Huang and May, J. Acoust. Soc. Am. (in press)]. The present study investigates how directional cues conveyed by the mid- and high-frequency spectrum of the HRTF influence the cat's ability to discriminate between sound locations. Thresholds for spatial acuity were measured as minimum audible angles (MAAs) [Mills, J. Acoust. Soc. Am. 30, 237-246 (1958)] at positive azimuths in the interaural horizontal plane and at positive and negative elevations in the median vertical plane. The frequency domain of the noise burst had little effect on MAAs in the horizontal plane, but removal of high-frequency spectral information significantly increased thresholds at positive and negative elevations in the median plane. These results suggest that cats are sensitive to directional properties of the HRTF at frequencies above 18 kHz and may use this information to detect small changes in sound source elevation.

Animals↗

Spatial aspects of reproduced sound in small rooms.

This paper reports on the influence of individual reflections on the auditory spatial aspects of reproduced sound. The sound field produced by a single loudspeaker positioned in a normal listening room has been simulated using an electroacoustical synthesis of the direct sound, 17 individual reflections and the reverberant field. The threshold of detection was measured using the method of adjustment for five reflections using three subjects for noise and speech. The thresholds have been measured for two simulated situations (1) a loudspeaker with a frequency independent directivity characteristics and frequency independent absorption coefficients of the room surfaces and (2) a loudspeaker with directivity similar to a standard two-way system and absorption coefficients according to measurements of real materials. The results have shown that subjects can reliably distinguish between timbre and spatial aspect of the sound field, that the spectral energy above 2 kHz of the individual reflection determines the importance of the reflection for the spatial aspects, and that only the first order floor reflection will contribute to the spatial aspects.

Amplifiers, Electronic↗

Identification and localization of sound sources in the median sagittal plane.

The ability of human listeners to identify broadband noises differing in spectral structure was studied for multiple sound-source locations in the median sagittal plane. The purpose of the study was to understand how sound identification is affected by spectral variations caused by directionally dependent head-related transfer functions. It was found that listeners could accurately identify noises with different spectral peaks and valleys when the source location was fixed. Listeners could also identify noises when the source location was roved in the median sagittal plane when the relevant spectral features were at low frequency. Listeners failed to identify noises with roved location when the spectral structure was at high frequency, presumably because the spectral structure was confused with the spectral variations caused by different locations. Parallel experiments on sound localization showed that listeners can localize noises that they cannot identify. The combination of identification and localization experiments leads to the conclusion that listeners cannot compensate for directionally dependent filtering by their own heads when they try to identify sounds.

Adult↗

COURTSHIP SOUND PRODUCTION IN TWO SYMPATRIC SIBLING DROSOPHILA SPECIES.

Courting males of Drosophila persimilis produce a pulsed "vibration sound," with frequency of oscillation and pulse repetition rate markedly different from that of D. pseudoobscura. These differences probably contribute to reproductive isolation. Flight frequency is the same in both species. Brief "preliminary sounds" are common and the same in both species. Males with large parts of both wings removed produce nearly normal vibration sounds, and normal preliminary sounds.

Animals↗

A panoramic code for sound location by cortical neurons.

By conventional spike count measures, auditory neurons in the cat's anterior ectosylvian sulcus cortical area are broadly tuned for the location of a sound source. Nevertheless, an artificial neural network was trained to classify the temporal spike patterns of single neurons according to sound location. The spike patterns of 73 percent of single neurons coded sound location with more than twice the chance level of accuracy, and spike patterns consistently carried more information than spike counts alone. In contrast to neurons that are sharply tuned for location, these neurons appear to encode sound locations throughout 360 degrees of azimuth.

Acoustic Stimulation↗

Validity and reliability of acoustic analysis of respiratory sounds in infants.

OBJECTIVE: To investigate the validity and reliability of computerised acoustic analysis in the detection of abnormal respiratory noises in infants. METHODS: Blinded, prospective comparison of acoustic analysis with stethoscope examination. Validity and reliability of acoustic analysis were assessed by calculating the degree of observer agreement using the kappa statistic with 95% confidence intervals (CI). RESULTS: 102 infants under 18 months were recruited. Convergent validity for agreement between stethoscope examination and acoustic analysis was poor for wheeze (kappa = 0.07 (95% CI, -0.13 to 0.26)) and rattles (kappa = 0.11 (-0.05 to 0.27)) and fair for crackles (kappa = 0.36 (0.18 to 0.54)). Both the stethoscope and acoustic analysis distinguished well between sounds (discriminant validity). Agreement between observers for the presence of wheeze was poor for both stethoscope examination and acoustic analysis. Agreement for rattles was moderate for the stethoscope but poor for acoustic analysis. Agreement for crackles was moderate using both techniques. Within-observer reliability for all sounds using acoustic analysis was moderate to good. CONCLUSIONS: The stethoscope is unreliable for assessing respiratory sounds in infants. This has important implications for its use as a diagnostic tool for lung disorders in infants, and confirms that it cannot be used as a gold standard. Because of the unreliability of the stethoscope, the validity of acoustic analysis could not be demonstrated, although it could discriminate between sounds well and showed good within-observer reliability. For acoustic analysis, targeted training and the development of computerised pattern recognition systems may improve reliability so that it can be used in clinical practice.

Acoustics↗

Complete right bundle-branch block: echophonocardiographic study of first heart sound and right ventricular contraction times.

High speed enchocardiograms of the mitral, tricuspid, and pulmonary valves were recorded with a simultaneous electrocardiogram and phonocardiogram in 20 patients with complete right bundle-branch block and in 67 normal subjects. Late opening of the pulmonary valve indicating late right ventricular ejection was found in all patients. In 8 patients with wide splitting of the first heart sound the late ejection was related mainly to delay in tricuspid valve closure, suggesting a late onset of the right ventricular pressure pulse. In 10 patients with a single first heart sound the delayed ejection was associated with a long interval between tricuspid valve closure and pulmonary valve opening, suggesting a slow rising right ventricular pressure pulse; 3 of these patients also had late tricuspid valve closure but the tricuspid component of the first sound was absent. Late onset of pressure rise is thought to result from block in the main right bundle-branch, and a slow rising pulse from block in the distal Purkinje network. These findings explain the conflicting results in previous studies of the first heart sound and right ventricular pressure pulse in patients with right bundle-branch block, and may have prognostic significance.

Adolescent↗

Presystolic mitral closure sound in aortic regurgitation with left ventricular hypertrophy and first degree heart block.

We report a patient who developed aortic regurgitation and first degree atrioventricular block caused by infective endocarditis complicating aortic valve stenosis. There was premature closure of the mitral valve and, in the absence of an Austin Flint murmur, a simultaneous high frequency sound was audible which we regard as a presystolic first heart sound. That such a sound may be associated with valve closure, even though this precedes electrical and mechanical ventricular systole, provides further support for the valvular origin of the first heart sound.

Aged↗

Neural mechanisms for sound localization.

Although the efforts to find a place map of sound direction within the auditory system of mammals has been reinspired by the recent discoveries in owl, progress to date has not been encouraging. Neither the inferior colliculus nor auditory cortex has yielded immediate evidence of such a map, despite ingenious and persistent efforts to find it. Thus, at present, the evidence suggests that a head-referenced map of auditory space is more likely to be found in structures more motor than sensory in function--in the deep layers of the superior colliculus or brainstem tegmentum, for example. Insofar as these structures have been implicated in eye, ear, and head orientation toward a sound source, one might expect that premotor units for orienting would be sensitive to sound direction and thus, collectively, constitute a map of auditory azimuth isomorphic to the map of motor azimuth. However, even for these structures, the possibility for significant variation among mammalian species exists. Because many candidate motor structures (such as the deep superior colliculus) receive input from the cerebral cortex, and because the role of auditory cortex in sound localization seems to vary widely among mammals (38, 51) an equal amount of variation in auditory-motor maps may also exist.

Animals↗

Sound localization by human listeners.

In keeping with our promise earlier in this review, we summarize here the process by which we believe spatial cues are used for localizing a sound source in a free-field listening situation. We believe it entails two parallel processes: 1. The azimuth of the source is determined using differences in interaural time or interaural intensity, whichever is present. Wightman and colleagues (1989) believe the low-frequency temporal information is dominant if both are present. 2. The elevation of the source is determined from spectral shape cues. The received sound spectrum, as modified by the pinna, is in effect compared with a stored set of directional transfer functions. These are actually the spectra of a nearly flat source heard at various elevations. The elevation that corresponds to the best-matching transfer function is selected as the locus of the sound. Pinnae are similar enough between people that certain general rules (e.g. Blauert's boosted bands or Butler's covert peaks) can describe this process. Head motion is probably not a critical part of the localization process, except in cases where time permits a very detailed assessment of location, in which case one tries to localize the source by turning the head toward the putative location. Sound localization is only moderately more precise when the listener points directly toward the source. The process is not analogous to localizing a visual source on the fovea of the retina. Thus, head motion provides only a moderate increase in localization accuracy. Finally, current evidence does not support the view that auditory motion perception is anything more than detection of changes in static location over time.

Attention↗

Noninvasive method for detecting laryngeal narrowing with low-frequency sound.

We measured laryngeal narrowing with low-frequency sound in human subjects. A low-frequency sound of 800 Hz was forced into the mouth, and sound-pressure amplitude above (SPAa) and below the vocal cord (SPAb) was detected using two separate microphones at the anterior neck. If the subject voluntarily narrowed the larynx at functional residual capacity, the increased respiratory resistance (Rrs) was only caused by increased laryngeal resistance, and SPAa was increased and SPAb was decreased. The percent changes of SPAa (SPAa%) minus that SPAb (SPAb%) from the initial values (SPAa% - SPAb% = Y) was proportional to the increase of Rrs from the initial control state (X, cmH2O X 1-1 X s); Y = 22.4 X1.20 (coef of correlation, r = 0.96, P less than 0.01). We confirmed similar proportions in dogs by manually narrowing the vocal cord. When laryngeal resistance was directly measured by tracheal puncture with a needle, the proportions between Y and X were not significantly dependent on the increase of Rrs below the vocal cord induced by methacholine inhalation in human subjects and histamine injection in dogs. We concluded that the increase of laryngeal resistance from the initial state could be detected noninvasively by using low-frequency sound.

Animals↗

Transmission of sound generated by sternal percussion.

We indirectly determined the transmission path of sound generated by sternal percussion in five healthy subjects. We percussed the sternum of each subject while recording the output audio signal at the posterior left and right upper and lower lung zones. Sound measurements were done during apnea at functional residual capacity, total lung capacity, and residual volume both with the lungs filled with air and with an 80% He-20% O2 (heliox) gas mixture. Three acoustic indexes were calculated from the output sound pulse: the peak-to-peak amplitude, the peak frequency, and the mid-power frequency. We found that the average values of all indexes tended to be greater in the upper than in the ipsilateral lower lung zones. In the upper zones, peak-to-peak amplitude was greater at total lung capacity and residual volume than at functional residual capacity. Replacing air with heliox did not change these results. These experiments, together with others performed during Mueller and Valsalva maneuvers, suggest that resonance of the chest cage is the predominant factor determining the transmission of sternal percussion sounds to the posterior chest wall. The transmission seems to be only minimally affected by the acoustic characteristics of the lung parenchyma.

Adult↗

Transmission to the chest of sound introduced at the mouth.

We examined the transmission to the chest wall of white noise and 25-Hz square-wave-generated noise introduced at the mouth of five healthy subjects. The output audio signals were recorded over the left and right upper and lower lung zones, posteriorly. Sound measurements were made during apnea at functional residual capacity, total lung capacity, and residual volume both after breathing air and an 80% He-20% O2 (heliox) gas mixture. We calculated the peak-to-peak amplitude, the peak frequency, and the midpower frequency of the output sound. We found no consistent variations in the values of these indexes due to lung volume or resident gas density. In all cases, the transmitted sound was most intense at the right upper zone. This could not be explained on the basis of technical factors but was probably the result of normal asymmetry of the mediastinal anatomy. These data suggest that sound introduced through the mouth of healthy individuals excites intrathoracic structures but is transmitted through the parenchyma in such a manner that it is not markedly affected by familiar physiological variables. This must be taken into account if objective acoustical tests of lung physiology are to be developed.

Adult↗

Regional effects of myocardial ischemia on epicardially recorded canine first heart sounds.

To determine whether focal changes in myocardial material properties are important in determining the response of first heart sound acceleration amplitude and frequency to myocardial ischemia, cardiac vibrations were simultaneously recorded from ischemic and nonischemic regions of canine epicardium by use of ultralight acceleration transducers. Cardiac acceleration and hemodynamics were recorded before and 5 min, 15 min, 1 h, and 2 h after left circumflex coronary artery occlusion. Peak-to-peak amplitude declined transiently in the nonischemic zone during early occlusion (P < 0.05) but was not decreased at any time in the ischemic myocardium. The median frequency of first heart sound vibrations in the ischemic region increased 31% within 5 min after occlusion (P < 0.01) and remained elevated for 2 h (P < 0.05). Nonischemic zone frequency was not statistically different from baseline at any time point. The disparate regional response of first heart sound vibrational frequency to myocardial ischemia suggests that propagating mechanical transients and myocardial contractile acceleration, rather than resonant vibrations, produce the first heart sound.

Animals↗