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Choice and foraging.

In Experiment 1, six naive pigeons were trained on a foraging schedule characterized by different states beginning with a search state in which completion of a fixed-interval on a white key led to a choice state. In the choice state the subject could, by appropriate responding on a fixed ratio of three, either accept or reject the schedule of reinforcement that was offered (either a variable-interval five-second or a variable-interval 20-second). If the subject accepted the schedule, it entered a "handling state" in which the appropriate variable-interval schedule was presented. Completion of the variable-interval schedule produced food. The independent variable was the fixed-interval value in the search state, and the dependent variable was the rate of acceptance of the long variable-interval in the choice state. Experiment 2 was identical except that the search state required completion of a variable-interval, instead of a fixed-interval, schedule. The rate of acceptance of the long variable-interval schedule in both experiments was a direct function of the length of the search state, in accordance with both optimality theory and the delay-reduction hypothesis.

Journal Article↗

A temporal limit on the effect of future food on current performance in an analogue of foraging and welfare.

Rats obtained access to food twice each 24-hour period. The first session was a work session in which food was available on a progressive-ratio schedule. During the second session, which occurred between 1 and 23 hours after the work session, food was freely available up to a fixed total intake each 24 hours. The situation resembled elements of several real world circumstances, including the choice between continuing to forage in a rapidly depleting patch and waiting for a better patch, and between working now and receiving a guaranteed income later. The purpose of the experiment was to explore the time period over which future access to reward could affect current responding. Contrary to what might be expected from recent theorizing, anticipation of future food delayed by an hour or more after the start of the work session had no effect on current performance. Food intake was high and constant during work sessions except for a prefeeding effect that occurred when the free session closely preceded the next day's work session. Also, an increase in the difficulty of the work schedule increased the amount of work and the maximum price paid for food as if the work session were the only time food was available. The results indicate the importance of considering temporal limits in theories that require animals to integrate input over time to determine the allocation of resources among alternatives.

Journal Article↗

The matching law applies to wagtails' foraging in the wild.

Field data concerning the time budgets and foraging success of pied wagtails (Motacilla alba yarrelli, Gould) are reanalyzed. It is found that the data are well described by the generalized matching law, with a marked bias towards spending time on the territory. In this case matching is not the result of maximizing reward rate, but it remains possible that it results from an allocation of time that maximizes survival.

Journal Article↗

Choice, foraging, and reinforcer duration.

Pigeons were exposed to a foraging schedule characterized by three different states, beginning with a search state in which completion of a variable interval on a white key led to a choice state. In the choice state the subject could, by appropriate responding on a fixed ratio of three, either accept or reject the schedule offered. If the subject accepted the schedule, it entered a handling state in which the appropriate reinforcer amount was presented according to a variable-interval schedule. In Experiment 1 the shorter duration reinforcer was more likely to be accepted the longer the duration of the search state and the shorter the equal durations of the handling states. In Experiment 2 the shorter duration reinforcer was more likely to be accepted the longer the handling time preceding the longer duration reinforcer. All of the results were in qualitative--and some were in quantitative--agreement with those predicted by the delay-reduction hypothesis and the optimal-diet model.

Animals↗

Changes in feeding and foraging patterns as an antipredator defensive strategy: a laboratory simulation using aversive stimulation in a closed economy.

The effects of the risk of electric shock on the meal patterns of rats living in an operant chamber were investigated. Rats could obtain food by working on a response lever that provided reinforcement according to chained fixed-ratio continuous reinforcement schedules that allowed the animals control over meal size. Using a two-compartment operant chamber with a safe nesting area and manipulanda area with a grid floor, shock could be correlated with responding on the schedule. Shocks (less than or equal to 1.25 per hour) were scheduled to occur randomly throughout the day, independent of the rat's behavior. Shock caused a reorganization of meal patterns such that the animals took less frequent but larger meals. This pattern reduced the time the animals spent at risk without compromising caloric balance. Similar changes in feeding pattern were obtained in both hooded and albino rats. Exposure to shock in a separate chamber did not produce these behavioral modifications. The magnitude of shock-induced alterations of meal patterns was greater with chained fixed-ratio 90 continuous reinforcement than with chained fixed-ratio 10 continuous reinforcement. Additionally, the rats seemed to be able to reduce food intake but increase caloric efficiency, such that the reduced food intake did not have deleterious effects on maintenance of body weight. These behavioral modifications reduced the number of shocks received from that which would have been expected if meal pattern changes had not occurred. We suggest that this technique may provide a useful laboratory simulation of the impact that the risk of predation has on foraging behavior.

Animals↗

An eight-alternative concurrent schedule: foraging in a radial maze.

In two experiments conducted in an eight-arm radial maze, food pellets were delivered when a photocell beam was broken at the end of each arm via a nose poke, according to either fixed-interval or random-interval schedules of reinforcement, with each arm providing a different frequency of reinforcement. The behavior of rats exposed to these procedures was well described by the generalized matching law; that is, the relationships between log behavior ratios and log pellet ratios were approximated by linear functions. The slopes of these log-log functions, an index of sensitivity to reinforcement frequency, were greatest for nose pokes, intermediate for time spent in an arm, and least for arm entries. Similar results were obtained with both fixed-interval and random-interval schedules. Addition of a 10-s changeover delay in both experiments eliminated the slope differentials between nose pokes and time spent in an arm by reducing the slopes of the nose-poke functions. These results suggest that different aspects of foraging may be differentially sensitive to reinforcement frequency. With concurrent fixed-interval schedules, the degree of temporal control exerted by individual fixed-interval schedules was directly related to reinforcement frequency.

Animals↗

Automatic procedure exploiting multicommutation in flow analysis for simultaneous spectrophotometric determination of nonstructural carbohydrates and reducing sugar in forage materials.

A multicommutated flow procedure for simultaneous spectrophotometric determination of nonstructural carbohydrates and reducing sugar in forage materials is proposed. Determination of both analytes was based on the neucroine method after acid hydrolysis of nonstructural carbohydrates. Results for both analytes presented the following features: no significant difference at 90% confidence level when compared with reference method, linear response between 0.2 to 0.8% (w/v) (r = 0.999); relative standard deviations of < 2.0% (n = 10), and sampling rate of 32 determinations per hour.

Automation↗

Nitrogen and phosphorus status of soils and trophic state of lakes associated with forage-based beef cattle operations in Florida.

Forage-based livestock systems have been implicated as major contributors to deteriorating water quality, particularly for phosphorus (P) from commercial fertilizers and manures affecting surface and ground water quality. Little information exists regarding possible magnitudes of nutrient losses from pastures that are managed for both grazing and hay production and how these might impact adjacent bodies of water. We examined the changes that have occurred in soil fertility levels of rhizoma peanut (Arachis glabrata Benth.)-based beef cattle pastures (n = 4) in Florida from 1988 to 2002. These pastures were managed for grazing in spring followed by haying in late summer and were fertilized annually with P (39 kg P2O5 ha(-1)) and K (68 kg K2O ha(-1)). Additionally, we investigated trends in water quality parameters and trophic state index (TSI) of lakes (n = 3) associated with beef cattle operations from 1993 to 2002. Overall, there was no spatial or temporal buildup of soil P and other crop nutrients despite the annual application of fertilizers and daily in-field loading of animal waste. In fact, soil fertility levels showed a declining trend for crop nutrient levels, especially soil P (y = 146.57 - 8.14 x year; r2= 0.75), even though the fields had a history of P fertilization and the cattle were rotated into the legume fields. Our results indicate that when nutrients are not applied in excess, cow-calf systems are slight exporters of P, K, Ca, and Mg through removal of cut hay. Water quality in lakes associated with cattle production was "good" (30-46 TSI) based on the Florida Water Quality Standard. These findings indicate that properly managed livestock operations may not be major contributors to excess loads of nutrients (especially P) in surface water.

Agriculture↗

Mefluidide treatment of tall fescue pastures: forage quality.

Spring application of a plant growth regulator, mefluidide, to tall fescue (Festuca arundinacea Schreb.) pastures grazed from May to September reduced (P less than .05) available DM and digestible OM per hectare by 51 and 41%, respectively, compared to untreated pastures. However, seasonal averages for N, Ca and P content and in vitro OM disappearance (OMD) were greater (P less than .05) and NDF was lower (P less than .05) in tall fescue from mefluidide-treated pastures than in untreated pastures. Compared to hand-clipped samples, esophageal extrusa of tall fescue was 63% higher (P less than .05) in N (2.7 vs 1.7%) and 26% lower (P less than .05) in NDF (52.6 vs 69.9%) than hand-clipped tall fescue samples, regardless of treatment. Extrusa and hand-clipped herbage samples were similar (P greater than .10) in OMD during May and September, but OMD of extrusa was greater (P less than .05) during June, July and August compared to clipped samples. As a consequence of preventing stem elongation and seedhead formation earlier in the growing season, mefluidide treatment of tall fescue maintained forage quality at a higher level during midsummer.

Acetamides↗

Influence of forage diets on ruminal particle size, passage of digesta, feed intake and digestibility by steers.

To assess the influence of forage diets on particle size of digesta in the rumen, three ruminally fistulated steers were fed alfalfa, orchardgrass or switchgrass hays in a 3 x 3 Latin square design. Mean sieve size of ruminal digesta for alfalfa, orchardgrass and switchgrass diets was 671, 652 and 466 microns, respectively. A second experiment examined the influence of ruminal particle size on digesta passage, digestibility and intake. Four ruminally fistulated Angus-Hereford steers were given ad libitum access to different proportions of alfalfa and switchgrass in a 4 x 4 Latin square design. Either a 100% alfalfa, 50% alfalfa:50% switchgrass, 25% alfalfa:75% switchgrass or 100% switchgrass diet was offered once daily. Mean sieve sizes of the ruminal digesta of these diets responded cubically (P less than .01; 1,066, 946, 1,003 and 925 microns, respectively). Mean ruminal turnover times were 24.3, 24.8, 24.7 and 29.8 h, respectively. Dry matter intake increased linearly (P less than .10) as the proportion of legume in the diet increased; no influence of diet on DM digestibility was observed. Passage rate of dosed nylon particles 1, 3 and 5 mm in length was influenced (P less than .01) by size; however, no difference in the passage rates of the nylon particles due to diet was evident. Particle size of ruminal digesta did not respond in a linear manner to the proportion of legume in the diet. A smaller ruminal particle size was not associated with a faster turnover of digesta.

Animal Feed↗

Value of a molasses mixture containing natural protein as a supplement to brood cows offered low-quality forages.

In a 4-yr study, molasses (M; 6% CP), molasses-urea (MU; 17.5% CP; 92% molasses, 4% urea and 4% water) and molasses-cottonseed meal-urea (MCSM; 17.5% CP; 73% molasses, 25% cottonseed meal, 1% urea and 1% water) were compared as winter supplements for Braford cows (514 matings) consuming low-quality forage. Cows were fed 1.35, 1.50 and 1.27 kg.head-1.d-1 of the respective supplements for 127 d beginning in December. Cows were wintered on bahiagrass pasture (avg 8% CP) and stargrass hay (avg 5% CP). Quantity of hay offered averaged 7.6 kg.cow-1.d-1 over 109 d. Cows were exposed to bulls for 90 d beginning on March 1. Supplementation treatment did not affect cow live weight, live weight change or condition score (P greater than .05). However, pregnancy rate was higher (P less than .05) for cows fed MCSM (79.0%) than for cows fed M (67.8%). The pregnancy rate of cows fed MU (76.3%) was intermediate to, but not different (P greater than .05) from, that of cows fed either M or MCSM. Supplementation treatment did not affect calf performance as measured by birth weight, survival rate or weaning weight (P greater than .05). However, calves from cows fed MCSM were 6 d older (P less than .05) at weaning than calves from cows fed M. For 3-yr-old cows, pregnancy rate was considerably higher (P less than .05) for cows fed MCSM (69.6%) than for cows fed M (37.5%).(ABSTRACT TRUNCATED AT 250 WORDS)

Animal Feed↗

Effects of corn vs corn gluten feed on site, extent and ruminal rate of forage digestion and on rate and efficiency of gain.

Three digestion experiments and one growth experiment were conducted to determine site, extent and ruminal rate of forage digestion and rate and efficiency of gain by cattle offered alfalfa haylage supplemented with corn or dry corn gluten feed (CGF). In Exp. 1, eight steers were fed alfalfa haylage-based diets with substitution of corn for 0, 20, 40 or 60% of haylage in a 4 X 4 latin square. Increasing dietary corn substitution increased (P less than .05) OM, NDF and ADF digestion by steers but decreased (P less than .05) rate of in situ alfalfa DM digestion. In Exp. 2, five heifers were fed alfalfa haylage-based diets with increasing dietary levels of CGF in a 5 X 5 latin square. Increasing dietary CGF increased (P less than .05) OM, NDF and ADF digestion by heifers. In Exp. 3 and 4, cattle were fed alfalfa haylage-based diets containing either 20 or 60% corn or CGF. In Exp. 3, supplementation increased (P less than .05) OM and NDF digestion but level X supplement source interaction (P less than .05) occurred, with added CGF increasing OM and NDF digestion more than added corn. In Exp. 4, supplementation improved (P less than .05) DM intake, daily gain and feed efficiency. Dry matter intake and daily gain were greater (P less than .05) for 60% supplementation than for 20% supplementation. Overall, whereas increasing the level of dietary supplement increased (P less than .05) OM, NDF and ADF digestion, only corn addition decreased (P less than .05) rate of in situ alfalfa DM digestion. Daily gains and feed efficiencies were similar in cattle fed either corn or CGF with alfalfa haylage.

Animal Feed↗

Forage intake and nutrition of range goats in a shrubland in northeastern Mexico.

Voluntary intake and nutritive value of diets selected by goats grazing a shrubland at Marin county, N.L., Mexico were determined. Four esophageally cannulated Spanish female goats (average BW 35 kg), were used to collect extrusa samples monthly throughout 1 yr (June 1987 to May 1988). Organic matter intake (OMI) was estimated by total fecal collections from four castrated Spanish male goats (average BW 42 kg). Diets selected by goats were high in CP throughout the year (means = 18.9%). Available CP represented only about half the total CP. The ADF content of extrusa samples differed (P less than .05) among months, with high ADF values during winter months. Dietary lignin followed a similar pattern during the year. Goats consumed forage with sufficient amounts of Ca, Na, K, Mg, Cu, ZN, Mn, and Fe to meet requirements. The OMI of goats was different (P less than .05) among sampling periods. On the average, estimated OMI (means = 878.7 g/d) compared well with results of other studies in the area. Amounts of DE consumed by goats (1.1 Mcal DE/d) were not sufficient to meet requirements for maintenance plus low activity. High levels of browse in the diets presumably were responsible for low in vitro OM digestibility (means = 34.1%) throughout the year. Thus, productivity of range goats in these areas should be improved by supplementing with protein and energy.

Animal Feed↗

Effects of lactational status on forage intake, digestibility, and particulate passage rate of beef cows supplemented with soybean meal, wheat middlings, and corn and soybean meal.

Sixteen mature, lactating (453 kg) and 16 nonlactating (487 kg) Hereford and Angus x Hereford cows were used to determine effects of different dietary supplements and lactational status on forage intake, digestibility, and particulate passage rate. Supplement treatments and amounts fed (kg/d) were as follows: control, 0; and equal daily amounts of CP from soybean meal (SBM), 1.36; wheat middlings (WM), 3.41; or a blend of corn and soybean meal (corn-SBM; 22% corn and 76% SBM), 3.41. Cows were fed supplements at 0800 and had ad libitum access to prairie hay (4.9% CP) in stalls from 0800 to 1100 and from 1300 to 1600 for three 17-d periods. Lactational status and supplement type did not interact (P greater than .50) for hay DMI, DM digestibility, or particulate passage rate. Cows fed SBM ate more hay DM (P less than .01) and had greater hay DM digestibility (P less than .01) than did cows in other treatment groups. Average hay DMI (kg/100 kg of BW) was 1.95, 2.16, 1.94, and 1.89, and hay DM digestibility was 52, 61, 55, and 53% for control, SBM, WM, and corn-SBM supplements, respectively. Total diet DM digestibility was increased by supplementation (P less than .01), but no differences (P greater than .18) were observed among supplements. Lactating cows ate more (P = .13) hay DM (2.11 vs 1.87 kg/100 kg of BW) and had greater (P less than .05) fecal output (4.6 vs 4.3 kg/d) than did nonlactating cows. Dry matter digestibility and particulate passage rate were not affected (P greater than .35) by lactational status.(ABSTRACT TRUNCATED AT 250 WORDS)

Animal Feed↗

Effect of coliform bacteria, feed deprivation, and pH on ruminal D-lactic acid production by steer or continuous-culture microbial populations changed from forage to concentrates.

Fecal coliform bacteria were isolated from three herbivores (cattle, horse, and red panda) and shown to produce primarily the D-form of lactate, plus acetate and ethanol when grown anaerobically in 1.0% glucose broth. To evaluate coliform contribution to D-lactate acidosis in cattle, experiments involving a forage-adapted steer (fasted or normally fed) and four 500-ml fermentors were compared during 3 d of grain overload. In both systems, coliforms and D- and L-lactic acid production were greater from fasted than from normally fed steer inoculum. With fasted inoculum, coliform counts peaked (3 x 10(7)/ml at 7 h after initial engorgement) and receded to 10(3)/ml by the time D-lactate concentration peaked, indicating that bacteria other than coliform were responsible for the delayed peaking of D- (48 h) compared with L-lactate (24 h). Increases in lactobacilli more closely mimicked D-lactate increases than did changes in coliforms. The comparisons between the steer and fermentors showed many similar shifts in end-products and groups of bacteria, more so with the experiment initiated with fasted than with normal inoculum. With normal inoculum, VFA content and moles of butyrate/100 mol of VFA were greater in vitro than in vivo; VFA content presumably was larger because of VFA absorption in vivo. In a separate experiment, cultures initiated with identical inoculum and given the same amount of feed accumulated more lactate when pH was permitted to decrease to 5.0 than when pH was maintained at 5.5 for 6.0 or above, indicating the role buffers can have in controlling acidosis during diet change to concentrates.

Animals↗

Evaluation of dairy food processing wash water solids as a protein source: I. Forage intake, animal performance, ruminal fermentation, and site of digestion in heifers fed medium-quality hay.

Twelve ruminally, duodenally, and ileally cannulated (average initial BW 313 +/- 20 kg) and 27 intact Hereford heifers (average initial BW 256 +/- 17 kg) were used in two experiments to evaluate dairy food wash water solids (WWS) as a protein source in medium-quality hay diets. Heifers received a basal diet of orchardgrass hay (7.4% CP) and were assigned to one of three supplement treatments: control (C; .9% CP), WWS (18.8% CP)-, and soybean meal (SBM; 19.1% CP)-based supplements (fed at 1.5 kg of DM/d). Supplements were formulated to have similar ME concentrations. Ruminal ammonia concentrations were greater (P less than .10) for WWS- and SBM-supplemented heifers than for C heifers at most sampling times. Moreover, WWS and SBM increased (P less than .10) total VFA (mM) and acetate (mol/100 mol) and lowered propionate (mol/100 mol) at several sampling times. Ruminal fluid volume (liters) was unchanged (P greater than .10) by treatment; however, fluid dilution and flow rate (liters/h) were less (P less than .10) in C heifers than in heifers fed SBM or WWS supplements. Wash water solids and SBM supplementation increased (P less than .10) OM, NDF, and ADF digestibilities compared with C heifers. Feeding WWS and SBM supplements increased BW at 84 d (P less than .10) compared with C-supplemented heifers. Forage intake at 54 and 84 d by heifers supplemented with SBM or WWS was greater (P less than .10) than by C heifers. Control-supplemented heifers had the least, WWS intermediate, and SBM the greatest ADG at 84 d (P less than .10; .14 vs .35 vs .48 kg/d, respectively). These data indicate that WWS may be used as a protein source without serious adverse effects in heifers consuming medium-quality hay for 84 d.

Ammonia↗

Supplementation of ammoniated wheat straw: performance and forage utilization characteristics in beef cattle receiving energy and protein supplements.

Two experiments were conducted to evaluate performance and forage utilization characteristics of beef cattle fed ammoniated wheat straw (AWS) unsupplemented except for minerals or supplemented with energy and protein. In Exp. 1, 194 crossbred beef cows (BW = 472 kg) in late gestation were allotted by weight, breed type, and age during two consecutive winters to 12 drylot pens (three pens/treatment) for a 60-d feeding trial. The AWS (3% NH3 wt/wt) was tubground through a 7.6-cm screen, and cows were allowed ad libitum access to the AWS. In Exp. 2, 16 ruminally fistulated Angus x Hereford steers (BW = 300 kg) were blocked by weight and randomized to treatments in a 35-d intake-digestion trial. Daily supplementation treatments in both experiments were Control, no supplemental energy or protein; LSG, 1.36 kg of sorghum grain (SG); HSG, 2.72 kg of SG; and SG + SBM, 1.02 kg of SG + .34 kg of soybean meal (SBM). All animals received .23 kg of mineral mixture formulated to meet a pregnant cow's mineral requirements. Supplements LSG and SG + SBM were fed to provide the same daily ME, and HSG and SG + SBM were fed to provide the same daily CP. Cows were managed as one group during and after calving. In Exp. 1, all supplements increased gain (P less than .01) vs Control, and cows fed SG + SBM had higher (P = .05) gains than those fed LSG. The SG + SBM supplement increased (P less than .01) change in cow body condition score compared with LSG.(ABSTRACT TRUNCATED AT 250 WORDS)

Animal Feed↗

Forage systems for beef production from conception to slaughter: II. Stocker systems.

Fall weaned Angus calves grazed stockpiled 1) tall fescue (Festuca arundinacea Schreb.), 2) tall fescue-red clover (Trifolium pratense L.), or 3) tall fescue-alfalfa (Medicago sativa L.) or were barn-fed, 4) tall fescue hay, 5) orchardgrass (Dactylis glomerata L.)-alfalfa hay, or 6) tall fescue silage from late October to early April during each of 5 yr. Infection of the fescue with Acremonium coenophialum ranged from 0 to 55%. There were two replications each of steers and heifers for each forage system in a completely random design. Each replicate was grazed by three Angus stockers, except for System 1, which was grazed by six stockers, for a total of 420 stockers. Each pasture replicate contained .8 ha (except System 1, which was 1.6 ha), and the stocking rate was one stocker per .27 ha. Fescue hay and silage were harvested each spring for barn-fed systems from the area stockpiled for grazing by cattle in System 1. Nitrogen fertilizer (90 kg/ha) was applied in early spring and again in early August, before stockpiling; no N was applied to stockpiled fescue grown with legumes. Daily gains by calves grazing stockpiled fescue-alfalfa were greater (P less than .01) than by calves grazing stockpiled fescue-red clover or N-fertilized stockpiled fescue (.50, .33, and .34 kg/d, respectively), but fescue-alfalfa calves required more days (P less than .01) of supplemental hay feeding (105, 60, and 36, respectively). Calves fed fescue hay in the barn gained more (P less than .01) than those fed fescue silage.(ABSTRACT TRUNCATED AT 250 WORDS)

Animal Feed↗