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Frequency- and level-dependent changes in auditory brainstem responses (ABRS) in developing mice.

The development of the auditory brainstem response was studied to quantitatively assess its dependence on stimulus frequency and level. Responses were not observed to stimuli > or =16 kHz on P12, however, the full range of responsive frequencies included in the study was observed by P14. Response thresholds were high on P12, exceeding 100 dB SPL for all stimuli tested. The rate of threshold development increased progressively for stimulus frequencies between -2 and 10 kHz, with the most rapid changes occurring at frequencies >10 kHz. Adultlike thresholds were observed by P18. Response latencies and interpeak intervals matured rapidly over the course of the second and third postnatal weeks and did not achieve adultlike characteristics until after P18. Latencies of higher-order peaks were progressively and sequentially delayed relative to wave I. Wave I amplitudes developed nonmonotonically, growing during the first 24 days and stabilizing at adult values by approximately P36. Slopes of wave I amplitude-and latency-level curves were significantly steeper than those of adults during the neonatal period and the outcome of input-output analyses, as well as frequency-specific maturational profiles, support developmental models in which function initially matures in the mid-frequency range and proceeds, simultaneously, in both apical and basal directions.

Aging↗

Spectral loudness summation for short and long signals as a function of level.

The equal-loudness-ratio hypothesis (ELRH) assumes that the loudness ratio between equal-level long and short signals with the same spectrum is independent of level and spectrum. Thus, it predicts the same amount of spectral loudness summation for equal-level short and long signals. To investigate how duration and level affect spectral loudness summation, loudness functions for 10- and 1000-ms bandpass-filtered noise centered at 2 kHz were obtained for normal-hearing listeners using categorical loudness scaling. The bandwidth was varied from 200 to 6400 Hz. From the loudness functions the level difference between the 3200-Hz-wide reference signal and the equally loud test signals was determined for reference levels in the range from 25 to 105 dB SPL. In addition, for two reference levels (45 and 65 dB SPL), this level difference was measured using a loudness-matching procedure. Both procedures showed similar results for the two reference levels. Over a wide level range spectral loudness summation is larger for the short than for the equal-level long signals. This is in contrast to the predictions of the ELRH. However, the data at moderate levels are consistent with the expectations of a modified ELRH which assumes a smaller loudness ratio for broadband than for narrow-band signals.

Adult↗

Directional loudness in an anechoic sound field, head-related transfer functions, and binaural summation.

The effect of sound incidence angle on loudness was investigated using real sound sources positioned in an anechoic chamber. Eight normal-hearing listeners produced loudness matches between a frontal reference location and seven sources placed at other directions, both in the horizontal and median planes. Matches were obtained via a two-interval, adaptive forced-choice (2AFC) procedure for three center frequencies (0.4, 1, and 5 kHz) and two overall levels (45 and 65 dB SPL). The results showed that loudness is not constant over sound incidence angles, with directional sensitivity varying over a range of up to 10 dB, exhibiting considerable frequency dependence, but only minor effects of overall level. The pattern of results varied substantially between subjects, but was largely accounted for by variations in individual head-related transfer functions. Modeling of binaural loudness based on the at-ear signals favored a sound-power summation model, according to which the maximum binaural gain is only 3 dB, over competing models based on larger gains, or on the summation of monaural loudness indices.

Adult↗

Effect of modulation maskers on the detection of second-order amplitude modulation with and without notched noise.

The mechanisms underlying the detection of second-order amplitude modulation (AM) were explored. The detectability of second-order AM (fixed depth for each subject) was measured for first- and second-order modulation rates of 16 and 2 Hz, respectively (slow-rate pair), and 50 and 10 Hz, respectively (fast-rate pair), with no masker, a low-band modulation masker (centered at 2 or 10 Hz), and a high-band modulation masker (centered at 16 or 50 Hz). This was done in the absence and presence of an audio-frequency notched noise centered at the carrier frequency of 4000 Hz. Both modulation maskers were "low-noise" noises, to prevent overmodulation. In the absence of notched noise, both modulation maskers impaired performance for the slow-rate pair, but only the low-band masker impaired performance for the fast-rate pair. When notched noise was present, the low-band masker had no significant effect for either rate pair and the high-band masker had an effect only for the slow-rate pair. These results suggest that second-order AM detection is mediated both by an envelope distortion component at the second-order rate and by slow fluctuations in the output of a modulation filter tuned to the first-order rate. When notched noise is present, the distortion component plays little role.

Adult↗

Monaural and binaural loudness of 5- and 200-ms tones in normal and impaired hearing.

The difference in level required to match monaural and binaural loudness of 5- and 200-ms tones was measured for listeners with normal and impaired hearing. Stimuli were 1-kHz tones presented at levels ranging from 10 to 90 dB sensation level. Sixteen listeners (eight normal and eight with losses of primarily cochlear origin) made loudness matches between equal-duration monaural and binaural tones using an adaptive 2AFC procedure. The present results corroborate existing data for 200-ms tones in normal listeners and provide new data for 5-ms tones. On average, the binaural level difference required for equal loudness of monaural and binaural tones is about the same for 5- and 200-ms tones of equal level and changes as a function of level. The group data for normal and impaired listeners are in reasonable agreement with data in the literature. However, the data from some of the impaired listeners deviate markedly from the average, indicating that group data do not accurately represent the behavior of all impaired listeners. Derived loudness functions from the loudness-matching data are reasonably consistent with individual data in the literature.

Acoustic Stimulation↗

The detection of increments and decrements is not facilitated by abrupt onsets or offsets.

Two experiments measured thresholds for the detection of increments and decrements in the intensity of a quasi-continuous broadband-noise (experiment 1) or increments in a 477-Hz pure-tone pedestal (experiment 2). A variety of onset and offset ramps for the intensity change were tested, from instantaneous onsets or offsets to ramps lasting several tens of milliseconds. For increments and decrements with equal duration, the characteristics of the ramps had little effect on performance. Abrupt rise times, which are associated with strong transient responses in auditory neurons, did not facilitate detection in comparison to much slower rise times. The temporal window model of temporal resolution provided a good account of the data when the decision statistic was the maximum magnitude of the change in the output of the window produced by the increment or decrement, but provided a poor account of the data when the decision statistic was the maximum rate of change in the output of the window over time. Overall the results suggest that, in the absence of cues in the audio-frequency domain, rapid changes in envelope contribute little to near-threshold increment or decrement detection.

Adolescent↗

Amplitude modulation sensitivity as a mechanism increment detection.

Detectability of a tonal signal added to a tonal masker increases with increasing duration ("temporal integration"), up to some maximum duration. Initially assumed to be some form of energy integration over time, this phenomenon is now often described as the result of a statistical "multiple looks" process. For continuous maskers, listeners may also use a mechanism sensitive to changes in stimulus intensity, possibly a result of inherent sensitivity to amplitude modulation (AM). In order to examine this hypothesis, change detection was investigated in the presence of AM maskers presented at either the same carrier frequency as the target signal or at a distant frequency. The results are compatible with the hypothesis that listeners detect intensity increments by using change-detection mechanisms (modeled here as the outputs of a bank of modulation filters) sensitive to envelope modulation at both low (4-16 Hz) and high (around 100 Hz) rates. AM masking occurred even when the masker was at a carrier frequency more than two octaves above that of the signal to be detected. This finding is also compatible with the hypothesis that similar mechanisms underlie sensitivity to AM (where across-frequency masking is commonly shown) and detection of intensity increments.

Acoustic Stimulation↗

No adaptation in the amplitude modulation domain in trained listeners.

The present study shows that on average, exposure to a 15 min, 5 kHz tone modulated sinusoidally in amplitude at 16 Hz with a 100% depth does not affect significantly amplitude modulation (AM) detection thresholds measured between 4 and 64 Hz when listeners are extensively trained to the AM detection task, with and without adaptor before data collection. These results are compatible with previous work given that a clear 6-dB adaptation effect was observed during the first pilot trials. However, the results reveal that adaptation effects are not robust, and suggest that the mechanisms underlying adaptation to AM must be reevaluated.

Acoustic Stimulation↗

Potential carry-over effect in the measurement of induced loudness reduction.

The majority of studies on induced loudness reduction (ILR) use an experimental paradigm that results in an underestimation of the amount of ILR. Most of those studies utilize loudness matches between tones of two different frequencies (a test tone and a comparison tone) with (experimental condition) and without (baseline condition) an inducer tone at the test frequency. The change in level of the comparison tone between the baseline and experimental conditions is the amount of ILR. In those experiments, the level of the comparison tone in the baseline condition tends to be substantially higher (often about 10 dB) than in the experimental condition. Because of this level difference, exposure to the baseline condition immediately prior to the experimental condition causes unintended ILR for the comparison tone. In this study, the delay between the baseline and experimental conditions was varied and it was determined that the amount of ILR is underestimated by about 30% and the variability is increased when the experimental condition is run immediately after the baseline condition. A second experiment using a Békésy-tracking procedure showed that ILR maximizes rapidly upon exposure to an inducer and decays over the course of several minutes after the inducer is removed.

Acoustic Stimulation↗

Effects of vocal loudness variation on spectrum balance as reflected by the alpha measure of long-term-average spectra of speech.

The overall slope of long-term-average spectrum (LTAS) decreases if vocal loudness increases. Therefore, changes of vocal loudness also affects the alpha measure, defined as the ratio of spectrum intensity above and below 1000 Hz. The effect on alpha of loudness variation was analyzed in 15 male and 16 female voices reading a text at different degrees of vocal loudness. The mean range of equivalent sound level (L(eq)) amounted to about 28 dB and the mean range of alpha to 19.0 and 11.7 dB for the female and male subjects. The L(eq) vs. alpha relationship could be approximated with a quadratic function, or by a linear equation, if softest phonation was excluded. Using such equations alpha was computed for all values of L(eq) observed for each subject and compared with observed values. The maximum and the mean absolute errors were 2.4 dB and between 0.1 and 0.6 dB. When softest phonation was disregarded and linear equations were used, the maximum error was less than 2 dB and the mean absolute errors were between 0.2 and 0.7 dB. The strong correlation between L(eq) and alpha indicates that for a voice L(eq) can be used for predicting alpha.

Adult↗

Cues for masked amplitude-modulation detection.

The ability of psychoacoustic models to predict listeners' performance depends on two key stages: preprocessing and the generation of a decision variable. The goal of the current study was to determine the perceptually relevant decision variables in masked amplitude-modulation detection tasks in which the modulation depth of the masker was systematically varied. Potential cues were made unreliable by roving the overall modulation depth from trial to trial or were reduced in salience by equalizing the envelope energy of the standard and target after the signal was added. Listeners' performance was significantly degraded in both paradigms compared to the baseline (fixed-level modulation masker) condition, which was similar to those used in previous studies of masking in the envelope-frequency domain. Although this observation was broadly consistent with a simple long-term envelope power-spectrum model, there were several aspects of the data that were not. For example, the steep rate of change in threshold with masker depth and the fact that an optimal amount of envelope noise could enhance performance were not predicted by decision variables calculated directly from the stimulus envelope. A physiologically based processing model suggested a realistic nonlinear mechanism that could give rise to these second-order features of the data.

Audiometry, Pure-Tone↗

Prediction of absolute thresholds and equal-loudness contours using a modified loudness model.

The loudness model described by Moore et al. [J. Audio Eng. Soc. 45, 224-240 (1997)] forms the basis for a recent ANSI standard for the calculation of the loudness of steady sounds. However, the model does not give accurate predictions of the absolute thresholds published in a recent ISO standard. Here it is described how the assumed middle-ear transfer function in the model can be modified to give more accurate absolute threshold predictions. The modified model also gives reasonably accurate predictions of the equal-loudness contours published in a recent ISO standard.

Auditory Threshold↗

Spectral-peak selection in spectral-shape discrimination by normal-hearing and hearing-impaired listeners.

Spectral-shape discrimination thresholds were measured in the presence and absence of noise to determine whether normal-hearing and hearing-impaired listeners rely primarily on spectral peaks in the excitation pattern when discriminating between stimuli with different spectral shapes. Standard stimuli were the sum of 2, 4, 6, 8, 10, 20, or 30 equal-amplitude tones with frequencies fixed between 200 and 4000 Hz. Signal stimuli were generated by increasing and decreasing the levels of every other standard component. The function relating the spectral-shape discrimination threshold to the number of components (N) showed an initial decrease in threshold with increasing N and then an increase in threshold when the number of components reached 10 and 6, for normal-hearing and hearing-impaired listeners, respectively. The presence of a 50-dB SPL/Hz noise led to a 1.7 dB increase in threshold for normal-hearing listeners and a 3.5 dB increase for hearing-impaired listeners. Multichannel modeling and the relatively small influence of noise suggest that both normal-hearing and hearing-impaired listeners rely on the peaks in the excitation pattern for spectral-shape discrimination. The greater influence of noise in the data from hearing-impaired listeners is attributed to a poorer representation of spectral peaks.

Acoustic Stimulation↗

Effect of adaptive psychophysical procedure on loudness matches.

Large variability in equal-loudness matches has been observed across studies. The purpose of the present study was to gain insight into the extent to which this variability results from differences in psychophysical procedures and/or differences among listeners. Four adaptive two-interval, two-alternatives-forced-choice procedures were used to obtain equal-loudness matches between 5- and 200-ms 1-kHz tones as a function of level for each of six normal listeners. The procedures differed primarily in the sequence in which the stimuli were presented. The variations tested were the ordering of stimuli by amplitude across blocks of trials (both increasing and decreasing amplitudes), randomizing the order across those blocks, and randomizing the order within blocks. The random-within-block procedure, which sought to randomize any intertrial information, yielded a significantly greater amount of temporal integration than the other three procedures. The results show significant differences in temporal integration measurements at moderate levels for the same listeners across different procedures. Therefore, although there are individual differences among listeners in the amount of temporal integration measured across paradigms, the choice of paradigm also affects the amount of temporal integration measured at moderate levels.

Acoustic Stimulation↗

On first rahmonic amplitude in the analysis of synthesized aperiodic voice signals.

Rahmonics comprise the prominent peaks in the cepstrum of voiced speech; their locations correspond to the fundamental period and its multiples. The amplitude of the first rahmonic, R1, has previously been used to indicate voice quality. Although a correspondence between R1 and the richness of the harmonic spectrum for voiced speech is well recognized, a formal description has remained absent. A theoretical description of rahmonic analysis of voiced speech containing aspiration noise is provided, leading to a characterization of R1. The theory suggests that R1 is directly proportional to the geometric mean harmonics-to-noise ratio (gmHNR), where the gmHNR is defined as the mean of the individual spectral (i.e., at specific frequency locations) harmonics-to-noise ratios in dB. This hypothesis is validated using synthetically generated voice signals. R1 is shown to be directly proportional to gmHNR (measured directly from the dB spectrum). It is shown that R1 (estimated from speech) is directly proportional to R1 taken from the glottal signal. R1 and gmHNR (measured spectrally) underestimate the actual gmHNR when (averaged) noise levels exceed harmonic levels. Limiting the number of harmonics in the analysis window overcomes this problem and also alleviates the (temporal) window length/f0 dependence of R1 when estimated period synchronously.

Algorithms↗

A test of the binaural equal-loudness-ratio hypothesis for tones.

It is well known that a tone presented binaurally is louder than the same tone presented monaurally. It is less clear how this loudness ratio changes as a function of level. The present experiment was designed to directly test the Binaural Equal-Loudness-Ratio hypothesis (BELRH), which states that the loudness ratio between equal-SPL monaural and binaural tones is independent of SPL. If true, the BELRH implies that monaural and binaural loudness functions are parallel when plotted on a log scale. Cross-modality matches between string length and loudness were used to directly measure binaural and monaural loudness functions for nine normal listeners. Stimuli were 1-kHz 200-ms tones ranging in level from 5 dB SL to 100 dB SPL. A two-way ANOVA showed significant effects of level and mode (binaural or monaural) on loudness, but no interaction between the level and mode. Consequently, no significant variations were found in the binaural-to-monaural loudness ratio across the range of levels tested. This finding supports the BELRH. In addition, the present data were found to closely match loudness functions derived from binaural level differences for equal loudness using the model proposed by Whilby et al. [J. Acoust. Soc. Am. 119, 3931-3939 (2006)].

Acoustics↗

Binaural summation of the loudness of pure tones.

Subjects made magnitude estimations of the loudness of pure tones (100, 400, and 1000 Hz), which were presented at equal and unequal sound pressure levels to the two ears. The results were consistent with a model of linear binaural loudness summation: The estimates of loudness approximated the linear sum of the loudness estimates of the individual left-ear and right-ear components. The relation between loudness and sound pressure, over moderate SPLs, can be described by a power function with an exponent of about 0.6 at 400 and 1000 Hz, 0.75 at 100 Hz.

Acoustic Stimulation↗

On the use of comfortable listening levels in speech experiments.

In order to investigate the effect of presentation subjects' labeling of speechlike sound patterns, synthetic stimuli were constructed, varying systematically FO contours, VOT, and F2 transitions. These stimuli were presented in random sequences at levels between 15 and 105 dB SPL to subjects with normal hearing. No significant response variation was observed in the range 40--100 dB SPL. Subjects' labeling behavior suddenly breaks down below levels of around 35 dB SPL. The secondary findings of the study are also discussed in terms of different specific processing strategies for different specific speech features.

Auditory Threshold↗