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PubMed1988

Central and peripheral representation of whispered and voiced speech.

Whispered speech is very different acoustically from normally voiced speech, yet listeners appear to have little trouble perceiving whispered speech. Two selective adaptation experiments explored the basis for the common perception of whispered and voiced speech, using two synthetic /ba/-/wa/ continua (one voiced, and one whispered). In the first experiment the endpoints of each series were used as adaptors, along with several nonspeech adaptors. Speech adaptors produced reliable labeling shifts of syllables matching in periodicity (i.e., whispered-whispered or voiced-voiced); somewhat smaller effects were found with mismatched periodicity. A periodic nonspeech tone with short rise time (the "pluck") produced adaptation effects like those for /ba/. These shifts occurred for whispered test syllables as well as voiced ones, indicating a common abstract level of representation for voiced and whispered stimuli. Experiment 2 replicated and extended Experiment 1, using same-ear and cross-ear adaptation conditions. There was perfect cross-ear transfer of the nonspeech adaptation effect, again implicating an abstract level of representation. The results support the existence of two levels of processing for complex acoustic signals. The commonality of whispered and voiced speech arises at the second, abstract level. Both this level, and the earlier, more directly acoustic level, are susceptible to adaptation effects.

Attention↗
PubMed1991

Slippery context effect and critical bands.

This article explored the slippery context effect: When Ss judge the loudness of tones that differ in sound frequency as well as intensity, stimulus context (relative intensity levels at the 2 frequencies) can strongly influence the levels that are judged equally loud. It is shown that the size of the slippery context effect depends on the frequency difference between the tones: Small frequency differences (less than a critical bandwidth) produced essentially no slippery effect; much larger differences produced substantial effects. These results are consistent with a model postulating the existence of a central attentional or preattentive "filter-like" process whose weighting coefficients represent the size of the absolute as opposed to the relative (contextual) component of loudness perception and judgment.

Adult↗
PubMed1992

The role of spectral and dynamic cues in imagery for musical timbre.

The musical quality of timbre is based on both spectral and dynamic acoustic cues. Four 2-part experiments examined whether these properties are represented in the mental image of a musical timbre. Experiment 1 established that imagery occurs for timbre variations within a single musical instrument, using plucked and bowed tones from a cello. Experiments 2 and 3 used synthetic stimuli that varied in either spectral or dynamic properties only, to investigate imagery with strict acoustic control over the stimuli. Experiment 4 explored whether the dimension of loudness is stored in an auditory image. Spectral properties appear to play a much larger role than dynamic properties in imagery for musical timbre.

Adult↗
PubMed1992

Perceptual interactions between musical pitch and timbre.

These experiments examined perceptual interactions between musical pitch and timbre. Experiment 1, through the use of the Garner classification tasks, found that pitch and timbre of isolated tones interact. Classification times showed interference from uncorrelated variation in the irrelevant attribute and facilitation from correlated variation; the effects were symmetrical. Experiments 2 and 3 examined how musical pitch and timbre function in longer sequences. In recognition memory tasks, a target tone always appeared in a fixed position in the sequences, and listeners were instructed to attend to either its pitch or its timbre. For successive tones, no interactions between timbre and pitch were found. That is, changing the pitches of context tones did not affect timbre recognition, and vice versa. The tendency to perceive pitch in relation to other context pitches was strong and unaffected by whether timbre was constant or varying. In contrast, the relative perception of timbre was weak and was found only when pitch was constant. These results suggest that timbre is perceived more in absolute than in relative terms. Perceptual implications for creating patterns in music with timbre variations are discussed.

Adult↗
PubMed1992

Musical duplex perception: perception of figurally good chords with subliminal distinguishing tones.

In a variant of duplex perception with speech, phoneme perception is maintained when distinguishing components are presented below intensities required for separate detection, forming the basis for the claim that a phonetic module takes precedence over nonspeech processing. This finding is replicated with music chords (C major and minor) created by mixing a piano fifth with a sinusoidal distinguishing tone (E or E flat). Individual threshold intensities for detecting E or E flat in the context of the fixed piano tones are established. Chord discrimination thresholds defined by distinguishing tone intensity were determined. Experiment 2 verified masked detection thresholds and subliminal chord identification for experienced musicians. Accurate chord perception was maintained at distinguishing tone intensities nearly 20 dB below the threshold for separate detection. Speech and music findings are argued to demonstrate general perceptual principles.

Adult↗
PubMed1993

Phonetic coherence in duplex perception: effects of acoustic differences and lexical status.

A modified version of duplex perception was used to investigate the nature of perceptual organization of speech. Ss were asked to identify stimuli in which a third-formant transition was presented to 1 ear and a full syllable (base) was presented to the other ear. Phonetic integration occurred even when the spectral composition or onset frequency of the isolated transition was varied relative to the base. However, when onset asynchronies were added to spectral differences, the isolated transition ceased to contribute to the syllable percept. Lexical status of the eventual phonetic percepts also influenced the phonetic integration of acoustic components into syllable percepts. These results suggest that combinations of low-level acoustic and phonetic differences between components and higher level lexical information interact to influence the grouping of acoustic components into phonetic percepts.

Adult↗
PubMed1994

Developmental changes in perception of nonnative vowel contrasts.

Discrimination of 2 German vowel contrasts was examined in English-learning infants of 6-8 and 10-12 months of age using a head turn procedure. The younger infants were better able than the older infants to discriminate the nonnative contrasts, but performance at 6-8 months was below levels that have been reported for nonnative consonant contrasts. A 2nd experiment using a habituation looking procedure showed that 4-month-old infants discriminated both German vowel contrasts, but the 6-month-olds could not. The findings are consistent with previous consonant work, revealing a shift from a language-general toward a language-specific pattern during the 1st year of life. However, that shift begins earlier in development for vowels than for consonants.

Attention↗
PubMed1996

Spectral regularity as a factor distinct from harmonic relations in auditory grouping.

When the regular spectral pattern formed by an odd-harmonic complex (the base) is disrupted by an added even harmonic, the added component is judged as more salient than its neighbors. This study considered whether the effects of spectral pattern on perceptual segregation are restricted to harmonic stimuli. Participants either rated the clarity or judged the relative pitch of a cued component in a series of complex tones. The difference in clarity between added and base components found for the harmonic complexes was not reduced when the complexes were made inharmonic either by a frequency shift or by spectral stretch or compression. However, the added-base difference could be abolished for an inharmonic complex when the distribution of components across frequency was made uniform. These findings suggest that spectral regularity is a factor distinct from harmonic relations in auditory grouping.

Attention↗
PubMed1996

The Doppler illusion: the influence of dynamic intensity change on perceived pitch.

Four studies illustrate a new auditory illusion associated with the Doppler effect and demonstrate a new influence of dynamic intensity change on perceived pitch. Experiment 1 confirmed the existence of a popular belief that the pitch of a moving sound source rises as the source approaches. Because there is no corresponding rise in frequency, the authors refer to the perceived pitch rise as the Doppler illusion. Experiment 2 confirmed that the effect occurs perceptually, so the belief in a "naive principle" of physics has a perceptual basis. Experiment 3 confirmed the effect does not occur under matched static conditions. Experiment 4 showed that the influence of dynamic intensity change on perceived pitch occurs outside the realm of Doppler stimuli. The findings support a dynamic dimensional interaction of pitch and loudness, with marked differences in the perception of pitch and loudness under static and dynamic conditions.

Adult↗
PubMed1999

Perceptual organization of complex auditory sequences: effect of number of simultaneous subsequences and frequency separation.

Previous findings on streaming are generalized to sequences composed of more than 2 subsequences. A new paradigm identified whether listeners perceive complex sequences as a single unit (integrative listening) or segregate them into 2 (or more) perceptual units (stream segregation). Listeners heard 2 complex sequences, each composed of 1, 2, 3, or 4 subsequences. Their task was to detect a temporal irregularity within 1 subsequence. In Experiment 1, the smallest frequency separation under which listeners were able to focus on 1 subsequence was unaffected by the number of co-occurring subsequences; nonfocused sounds were not perceptually organized into streams. In Experiment 2, detection improved progressively, not abruptly, as the frequency separation between subsequences increased from 0.25 to 6 auditory filters. The authors propose a model of perceptual organization of complex auditory sequences.

Adult↗
PubMed2001

Evidence for hierarchical categorization of coarticulated phonemes.

The reported research investigates how listeners recognize coarticulated phonemes. First, 2 data sets from experiments on the recognition of coarticulated phonemes published by D. H. Whalen (1989) are reanalyzed. The analyses indicate that listeners used categorization strategies involving a hierarchical dependency. Two new experiments are reported investigating the production and perception of fricative-vowel syllables. On the basis of measurements of acoustic cues on a large set of natural utterances, it was predicted that listeners would use categorization strategies involving a dependency of the fricative categorization on the perceived vowel. The predictions were tested in a perception experiment using a 2-dimensional synthetic fricative-vowel continuum. Model analyses of the results pooled across listeners confirmed the predictions. Individual analyses revealed some variability in the categorization dependencies used by different participants.

Female↗
PubMed1991

Echoic memory in the rat: effects of inspection time, retention interval, and the spectral composition of masking noise.

Memory for tones (1100 vs. 2330 Hz) was studied in 4 rats (Rattus norvegicus), as affected by the durations of both target tones (30 to 620 ms) and noise-filled retention intervals (0 to 480 ms). With a 0-ms delay, performance was near asymptotic with the 30-ms tone, but the memory of this brief tone suffered a massive decrement at retention intervals as brief as 60 ms; in contrast, memory for the 340-ms tone was stable for at least 240 ms. If the retention interval was filled by band-stop noise (with targets presented in the spectral gap), then the rat's memory for brief tones was superior to that obtained with the standard broad-band noise filler, and band-stop noise was better than a band-pass noise that had the tones embedded in the region of its spectral energy. These findings are consistent with the hypotheses that auditory memory in the rat consists of a transient sensorylike echoic store and a short-term store more resistant to the effects of retroactive interference.

Animals↗
PubMed1990

Behavioral and physiological responses to maternal separation in squirrel monkeys (Saimiri sciureus).

This study extends an examination of the behavioral and pituitary-adrenal responses of infant squirrel monkeys (Saimiri sciureus) separated from their mothers under different environmental conditions to another physiological system by measuring the metabolites of the central monoamines found in the cerebrospinal fluid (CSF). This study included spectrographic examination of the vocalizations emitted by the infant during separation. Infants were separated from their mothers for 24 hr under 3 conditions: Home, infant remained in its home cage after removal of mother; adjacent, infant was placed in a cage adjacent to its mother; and total, infant was totally isolated. The behavioral results indicated that the number of calls emitted differed with condition (adjacent greater than total greater than home), and the peak frequency of the calls and number of multiple calls was greatest in the total condition. Plasma cortisol elevations after separation differentiated the conditions of separation (total greater than adjacent greater than home greater than base). The elevations in the CSF catecholamine metabolites (3-methoxy-4-hydroxyphenylglycol and homovanillic acid) were also sensitive to the conditions of separation (total greater than adjacent greater than base). These results are discussed in the context of coping theory.

Animals↗
PubMed1990

Ultrasonic vocalizations by rat pups in the cold: an acoustic by-product of laryngeal braking?

Isolated rat pups respond to cold exposure physiologically by increasing metabolic heat production and behaviorally by emitting ultrasound. The relationship between these 2 responses was investigated by monitoring oxygen consumption, heat production by brown adipose tissue, respiratory rate, and ultrasound production during cold exposure in pups 10-12 days of age. All 3 physiological measures increased contemporaneously with the initiation of ultrasound. Pups also exhibited a respiratory pattern characterized by the prolongation of expiratory duration in relation to inspiratory duration. Ultrasound was often detected during these prolonged expirations, suggesting that pups were using laryngeal braking. Laryngeal braking is thought to enhance oxygen uptake in the lungs. Thus, ultrasound may be an acoustic by-product of a respiratory maneuver that increases oxygen delivery to metabolically active tissues during cold exposure.

Adipose Tissue, Brown↗
PubMed1995

Pavlovian conditional vocalizations of the rat: a model system for analyzing the fear of pain.

Presentation of a 6-s light conditional stimulus (CS) that overlapped with a 1-s tailshock unconditional stimulus (US) generated audible conditional vocalization responses (VCRs) during the CS period. The rate of conditioning was observed to be directly related to the intensity of the tailshock US (0.15 mA-0.80 mA). The amplitude, duration, and number of VCRs was also directly related to US intensity, whereas the latency of VCRs from CS onset was inversely related to US intensity. VCRs were not observed in rats given explicitly unpaired presentations of CS and US (0.80 mA). The capacity of tailshock to support development of VCRs was found to depend on its capacity to elicit vocalization afterdischarges (VADs). Sonographic analysis of vocalizations revealed that VCRs and VADs share spectrographic characteristics. Results are discussed in terms of VCRs' providing a model system for analyzing the fear of pain and its suppression.

Animals↗
PubMed2000

Sexually stimulating signals of canary (Serinus canaria) songs: evidence for a female-specific auditory representation in the HVc nucleus during the breeding season.

During the breeding season under long-day conditions, male canaries sing sexually attractive songs and females respond behaviorally to such songs. This study assessed whether auditory response properties of neurons in nucleus HVc of female and male canaries are tuned to sexually salient song features: special song phrases and canary song segmentation. In sexually receptive female canaries, neurons responded to special song phrases with a decreased spike rate and were sensitive to canary song segmentation. The nonreceptive females showed no clear response to special song phrases. In females on short days, neurons responded to song phrases with an increase in activity. In males on long days, they exhibited phasic responses after the phrase onset, whatever the song phrase and song segmentation. This study demonstrates both a plasticity in relation to females' sexual responsiveness and a sexual dimorphism in the auditory processing performed in the HVc.

Animals↗
PubMed2000

Female presence during postejaculatory interval facilitates penile erection and 22-kHz vocalization in male rats.

During the postejaculatory interval (PEI), male rats exhibit prolonged immobility, 22-kHz vocalization, and penile erections. To test whether females modulate these behaviors, females were removed after the first or second ejaculation or left in the test chamber. Female presence during the PEI delayed exploratory behavior and facilitated vocalization and erection. Female stimulation of vocalization is consistent with the hypothesis that vocalization has a communicative function, not just a thermoregulatory one. The timing of the effect of females on erection suggests that males are sexually arousable well before they resume copulation. Therefore, erection may be better than vocalization as an indicator of the male's sexual refractoriness. The findings also challenge the conventional view that the PEI comprises absolute and relative sexual refractory periods marked, respectively, by the presence and absence of 22-kHz vocalization.

Animals↗
PubMed1994

On the perceptual organization of speech.

A general account of auditory perceptual organization has developed in the past 2 decades. It relies on primitive devices akin to the Gestalt principles of organization to assign sensory elements to probable groupings and invokes secondary schematic processes to confirm or to repair the possible organization. Although this conceptualization is intended to apply universally, the variety and arrangement of acoustic constituents of speech violate Gestalt principles at numerous junctures, cohering perceptually, nonetheless. The authors report 3 experiments on organization in phonetic perception, using sine wave synthesis to evade the Gestalt rules and the schematic processes alike. These findings falsify a general auditory account, showing that phonetic perceptual organization is achieved by specific sensitivity to the acoustic modulations characteristic of speech signals.

Adult↗